Size: 52 sq km (20 sq miles), Tanzania's smallest park.
Location: 16 km (10 miles) north of Kigoma on the shore of Lake Tanganyika in western Tanzania.

An excited whoop erupts from deep in the forest, boosted immediately by a dozen other voices, rising in volume and tempo and pitch to a frenzied shrieking crescendo. It is the famous ‘pant-hoot’ call: a bonding ritual that allows the participants to identify each other through their individual vocal stylisations. To the human listener, walking through the ancient forests of Gombe Stream, this spine-chilling outburst is also an indicator of imminent visual contact with man’s closest genetic relative: the chimpanzee.

Gombe is the smallest of Tanzania's national parks: a fragile strip of chimpanzee habitat straddling the steep slopes and river valleys that hem in the sandy northern shore of Lake Tanganyika. Its chimpanzees – habituated to human visitors – were made famous by the pioneering work of Jane Goodall, who in 1960 founded a behavioural research program that now stands as the longest-running study of its kind in the world. The matriarch Fifi, the last surviving member of the original community, only three-years old when Goodall first set foot in Gombe, is still regularly seen by visitors.

Chimpanzees share about 98% of their genes with humans, and no scientific expertise is required to distinguish between the individual repertoires of pants, hoots and screams that define the celebrities, the powerbrokers, and the supporting characters. Perhaps you will see a flicker of understanding when you look into a chimp's eyes, assessing you in return - a look of apparent recognition across the narrowest of species barriers.

The most visible of Gombe’s other mammals are also primates. A troop of beachcomber olive baboons, under study since the 1960s, is exceptionally habituated, while red-tailed and red colobus monkeys - the latter regularly hunted by chimps – stick to the forest canopy.

The park’s 200-odd bird species range from the iconic fish eagle to the jewel-like Peter’s twinspots that hop tamely around the visitors’ centre.

After dusk, a dazzling night sky is complemented by the lanterns of hundreds of small wooden boats, bobbing on the lake like a sprawling city.

Getting there
Kigoma is connected to Dar and Arusha by scheduled flights, to Dar and Mwanza by a slow rail service, to Mwanza, Dar and Mbeya by rough dirt roads, and to Mpulungu in Zambia by a weekly ferry.
From Kigoma, local lake-taxis take up to three hours to reach Gombe, or motorboats can be chartered, taking less than one hour.

What to do
Chimpanzee trekking; hiking, swimming and snorkelling. Visit the site of Henry Stanley's famous “Dr Livingstone I presume” at Ujiji near Kigoma, and watch the renowned dhow builders at work.

When to go
The chimps don't roam as far in the wet season (February-June, November-mid December) so may be easier to find;
better picture opportunities in the dry (July-October and late December).

Accommodation
1 new luxury tented lodge, as well a self-catering hostel, guest house and campsites on the lakeshore.

Size: 1,613 sq km (623 sq miles).

Location: Western Tanzania, bordering Lake Tanganyika.

Set deep in the heart of the African interior, inaccessible by road and only 100km (60 miles) south of where Stanley uttered that immortal greeting “Doctor Livingstone, I presume”, is a scene reminiscent of an Indian Ocean island beach idyll.

Silky white coves hem in the azure waters of Lake Tanganyika, overshadowed by a chain of wild, jungle-draped peaks towering almost 2km above the shore: the remote and mysterious Mahale Mountains.

Mahale Mountains, like its northerly neighbour Gombe Stream, is home to some of Africa’s last remaining wild chimpanzees: a population of roughly 800, habituated to human visitors by a Japanese research project founded in the 1960s. Tracking the chimps of Mahale is a magical experience. The guide's eyes pick out last night's nests - shadowy clumps high in a gallery of trees crowding the sky. Scraps of half-eaten fruit and fresh dung become valuable clues, leading deeper into the forest. Butterflies flit in the dappled sunlight.

Then suddenly you are in their midst: preening each other's glossy coats in concentrated huddles, squabbling noisily, or bounding into the trees to swing effortlessly between the vines.

The area is also known as Nkungwe, after the park's largest mountain, held sacred by the local Tongwe people, and at 2,460 metres (8,069 ft) the highest of the six prominent points that make up the Mahale Range.

And while chimpanzees are the star attraction, the slopes support a diverse forest fauna, including readily observed troops of red colobus, red-tailed and blue monkeys, and a kaleidoscopic array of colourful forest birds.

You can trace the Tongwe people's ancient pilgrimage to the mountain spirits, hiking through the montane rainforest belt – home to an endemic race of Angola colobus monkey - to high grassy ridges chequered with alpine bamboo. Then bathe in the impossibly clear waters of the world’s longest, second-deepest and least-polluted freshwater lake – harbouring an estimated 1,000 fish species - before returning as you came, by boat.

Getting there
Charter flight from Arusha, Dar or Kigoma.
Charter private or national park motorboat from Kigoma, three to four hours.
Weekly steamer from Kigoma, seven hours, then hire a local fishing boat or arrange with park HQ for pickup in park boat, another one or two hours.

What to do
Chimp tracking (allow two days); hiking; camping safaris; snorkelling; fish for your dinner.

When to go
Dry season (May-October) best for forest walks although no problem in the light rains of October/November.

Accommodation
Two seasonal luxury tented camps.
Two small resthouses, large campsite.

NOTE
The same rules for chimpanzee viewing at Gombe Stream apply at Mahale.

Size: 240 sq km (93 sq miles).

Location: Northwest Tanzania, 150 km (95 miles) west of Mwanza.

A pair of fish eagles guards the gentle bay, their distinctive black, white and chestnut feather pattern gleaming boldly in the morning sun. Suddenly, the birds toss back their heads in a piercing, evocative duet. On the sandbank below, a well-fed monster of a crocodile snaps to life, startled from its nap. It stampedes through the crunchy undergrowth, crashing into the water in front of the boat, invisible except for a pair of sentry-post eyes that peek menacingly above the surface to monitor our movements.

Rubondo Island is tucked in the southwest corner of Lake Victoria, the world's second-largest lake, an inland sea sprawling between Tanzania, Uganda and Kenya. With nine smaller islands under its wing, Rubondo protects precious fish breeding grounds.

Tasty tilapia form the staple diet of the yellow-spotted otters that frolic in the island’s rocky coves, while rapacious Nile perch, some weighing more than 100kg, tempt recreational game fishermen seeking world record catches.

Rubondo is more than a water wonderland. Deserted sandy beaches nestle against a cloak of virgin forest, where dappled bushbuck move fleet yet silent through a maze of tamarinds, wild palms, and sycamore figs strung with a cage of trailing taproots.

The shaggy-coated aquatic sitatunga, elsewhere the most elusive of antelopes, is remarkably easily observed, not only in the papyrus swamps it normally inhabits, but also in the forest interior.
Birds are everywhere.

Flocks of African grey parrots – released onto the island after they were confiscated from illegal exporters – screech in comic discord as they flap furiously between the trees.

The azure brilliance of a malachite kingfisher perched low on the reeds competes with the glamorous, flowing tail of a paradise flycatcher as it flits through the lakeshore forest. Herons, storks and spoonbills proliferate in the swampy lake fringes, supplemented by thousands of Eurasian migrants during the northern winter.

Wild jasmine, 40 different orchids and a smorgasbord of sweet, indefinable smells emanate from the forest.

Ninety percent of the park is humid forest; the remainder ranges from open grassland to lakeside papyrus beds.

A number of indigenous mammal species - hippo, vervet monkey, genet and mongoose - share their protected habitat with introduced species such as chimpanzee, black-and-white colobus, elephant and giraffe, all of which benefit from Rubondo's inaccessibility.

Getting there
Scheduled flights from Arusha, Lake Manyara, Serengeti and Mwanza in peak season, charter flights only in low season.
By road from Mwanza and then boat transfer. Contact the Park for transport details.

What to do
Walking safaris, boat excursions, sport fishing, chimpanzee treks, plans for canoe trips.

When to go
Dry season, June-August. Wildflowers and butterflies
Wet season November-March. December- February best for migratory birds.

Accommodation
One luxury tented camp, park bandas and campsite.

Size: 412.9 sq km (159 sq miles)

Location: Southern Tanzania.
The temporary park headquarters at Matamba are situated approximately 100km (60 miles) from Mbeya town.

Locals refer to the Kitulo Plateau as Bustani ya Mungu - The Garden of God – while botanists have dubbed it the Serengeti of Flowers, host to ‘one of the great floral spectacles of the world’. And Kitulo is indeed a rare botanical marvel, home to a full 350 species of vascular plants, including 45 varieties of terrestrial orchid, which erupt into a riotous wildflower display of breathtaking scale and diversity during the main rainy season of late November to April.

Perched at around 2,600 metres (8,500 ft) between the rugged peaks of the Kipengere, Poroto and Livingstone Mountains, the well-watered volcanic soils of Kitulo support the largest and most important montane grassland community in Tanzania.

One of the most important watersheds for the Great Ruaha River, Kitulo is well known for its floral significance – not only a multitude of orchids, but also the stunning yellow-orange red-hot poker and a variety of aloes, proteas, geraniums, giant lobelias, lilies and aster daisies, of which more than 30 species are endemic to southern Tanzania.
Big game is sparsely represented, though a few hardy mountain reedbuck and eland still roam the open grassland.

But Kitulo – a botanist and hiker’s paradise - is also highly alluring to birdwatchers. Tanzania’s only population of the rare Denham’s bustard is resident, alongside a breeding colony of the endangered blue swallow and such range-restricted species as mountain marsh widow, Njombe cisticola and Kipengere seedeater. Endemic species of butterfly, chameleon, lizard and frog further enhance the biological wealth of God’s Garden.

Getting there
4WD only.
From Chimala, 78km east of Mbeya along the surfaced main road to Dar es Salaam, head south along the rough but spectacular dirt road - called Hamsini na Saba (57) after the number of hairpin bends along its length - to the temporary park headquarters at Matamba, from where it’s another hour’s drive to the plateau.
Basic and erratic public transport is available.

What to do
Good hiking trails exist and will soon be developed into a formal trail system.
Open walking across the grasslands to watch birds and wildflowers.
Hill climbing on the neighbouring ranges. A half-day hike from the park across the Livingstone Mountains leads to the sumptuous Matema Beach on Lake Nyasa.

When to go
Wildflower displays peak between December and April.
The sunnier months of September to November are more comfortable for hiking but less rewarding to botanists.
Conditions are cold and foggy from June to August.

Accommodation
Mbeya is serviced by everything from luxury hotels to simple guesthouses, while two simple guesthouses also exist in Matamba.
There is no accommodation in the park as yet, but very basic accommodation and meals are available at the adjacent Kitulo Farm.
Three special campsites are planned within the park, catering to fully equipped campers.
Two moderately priced church-run hostels are situated on Matema Beach.